# 📐  L2 Spec — Wright-Fisher Population Genetics — Nominal + Mismatch Specs

**ID:** `L2-408-001` · **Status:** ⊙ Testnet (genesis catalog)
**Parents:** ⬆ Principle [`L1-408`](/artifact/l1/L1-408)

> **🎯 Tolerance (ε):** expression
> **⚖ Difficulty score:** 0.38 · **Delta tier:** 5
> **⛓ On-chain block:** 41555222

---

## 1. Domain Ω

**Type:** `range`

| Parameter | Unit | Range |
|---|---|---|
| S selection | — | -0.1 – 0.1 |
| Ne effective | — | 10 – 100000 |
| N generations | — | 5 – 100 |
| N sampled alleles | — | 10 – 1000 |

## 2. Tolerance (ε)

**Type:** `fn`

```
0.2 + 0.1 * (20 / N_generations) ** 0.5
```

## 3. Difficulty

| Property | Value |
|---|---|
| Difficulty score | 0.38 |
| Delta tier | 5 |

## 4. Fingerprint

`158b4fe2cdf341bcf7d624c60acef0bbd24b47e7113e9ddc2596ae8083eb4384`

## 5. On-chain Registration

- **Chain hash:** `0x9ea335280d1dbb3174ac9473ebb6575b4ab90570db5d63110a221e72844b788a`
- **Chain tx hash:** `0x9136ed4a11677ae5bb91d1b6310c39fba9e7974755a4dd03909e057f3db3aa6a`
- **Chain block:** `41555222`

---

## File Mapping

This bundle consists of: `L2-408-001.md`, `L2-408-001.json`.

| File | Role | How to regenerate |
|------|------|-------------------|
| `L2-408-001.md` | Source of truth — edit this | Human or LLM |
| `L2-408-001.json` | Structured metadata for the registry | LLM regenerates from the sections above |

**Prompt for your LLM after editing this Markdown:**

> Read the attached Markdown. Regenerate the sibling `.json` so every field matches.
> Preserve the schema documented in the rows above.
> Output each file in its own fenced code block tagged with the filename.
> Output only the JSON object.

_This Markdown was auto-synthesized from the catalog row for `L2-408-001`._
_Edit it, regenerate the JSON, and submit at [/submit](/submit) to claim the artifact._